K Moyes, BJT Morgan, A Morris, SJ Morris, TH Clutton-Brock, T Coulson (2009) Exploring individual quality in a wild population of red deer. Journal of Animal Ecology 78: 406-413 [PDF]

DH Nussey, AJ Wilson, A Morris, J Pemberton, T Clutton-Brock, LEB Kruuk (2008) Testing for genetic trade-offs between early- and late-life reproduction in a wild red deer population. Proceedings of the Royal Society B-Biological Sciences 275: 745-750 [PDF]

KV Stopher, JM Pemberton,TH Clutton-Brock, T Coulson (2008) Individual differences, density dependence and offspring birth traits in a population of red deer. Proceedings of the Royal Society B-Biological Sciences 275: 2137-2145 [PDF]

T Coulson, S Tuljapurkar (2008) The dynamics of a quantitative trait in an age-structured population living in a variable environment. American Naturalist 172: 599-612. [PDF]

PD McLoughlin, T Coulson, T Clutton-Brock (2008) Cross-generational effects of habitat and density on life history in red deer. Ecology 89: 3317-3326 [PDF]

DH Nussey, LEB Kruuk, A Morris,TH Clutton-Brock (2007) Environmental conditions in early life influence ageing rates in a wild population of red deer. Current Biology 17: R1000-R1001 [PDF]

AJ Wilson,  DH Nussey, JM Pemberton, JG Pilkington, A Morris, F Pelletier, TH Clutton-Brock, LEB Kruuk (2007) Evidence for a genetic basis of aging in two wild vertebrate populations. Current Biology 17: 2136-2142 [PDF]

K Foerster, T Coulson, BC Sheldon, JM Pemberton, TH Clutton-Brock, LEB Kruuk (2007) Sexually antagonistic genetic variation for fitness in red deer. Nature, 447: 1107-1110. [PDF]

J Hone, TH Clutton-Brock (2007) Climate, food, density and wildlife population growth rate. Journal of Animal Ecology, 76: 361-367. [PDF]

ST Kalinowski, ML Taper, TC Marshall (2007) Revising how the computer program CERVUS accommodates genotyping error increases success in paternity assignment. Molecular Ecology, 16: 1099-1106. [PDF]

LEB Kruuk, JD Hadfield (2007) How to separate genetic and environmental causes of similarity between relatives. Journal of Evolutionary Biology, 20: 1890-1903. [PDF]

M Sims, DA Elston, A Larkham, DH Nussey, SD Albon (2007) Identifying when weather influences life-history traits of grazing herbivores. Journal of Animal Ecology, 76: 761-770. [PDF]

DH Nussey, B Metherell, K Moyes, A Donald, FE Guinness, TH Clutton-Brock (2007) The relationship between tooth wear, habitat quality and late-life reproduction in a wild red deer population. Journal of Animal Ecology, 76: 402-412. [PDF]

J Slate, JM Pemberton (2007) Admixture and patterns of linkage disequilibrium in a free-living vertebrate population. Journal of Evolutionary Biology, 20: 1415-1427. [PDF]

K Csillery, T Johnson, D Beraldi, T Clutton-Brock, D Coltman, B Hansson, G Spong, JM Pemberton (2006) Performance of marker-based relatedness estimators in natural populations of outbred vertebrates. Genetics, 173: 2091-2101.  [PDF]

R Lande, S Engen, BE Saether, T Coulson (2006) Estimating density dependence from time series of population age structure. American Naturalist, 168: 76-87. [PDF]

PD McLoughlin, MS Boyce, T Coulson, T Clutton-Brock (2006) Lifetime reproductive success and density-dependent, multi-variable resource selection. Proceedings of the Royal Society B-Biological Sciences, 273: 1449-1454. [PDF]

K Moyes, T Coulson, BJT Morgan, A Donald, SJ Morris, TH Clutton-Brock (2006) Cumulative reproduction and survival costs in female red deer. Oikos, 115: 241-252. [PDF]

DH Nussey, LEB Kruuk, A Donald, M Fowlie, TH Clutton-Brock (2006) The rate of senescence in maternal performance increases with early-life fecundity in red deer. Ecology Letters, 9: 1342-1350. [PDF]

DH Nussey, J Pemberton, A Donald, LEB Kruuk (2006) Genetic consequences of human management in an introduced island population of red deer (Cervus elaphus). Heredity, 97: 56-65. [PDF]

RE Stevens, AM Lister, REM Hedges (2006) Predicting diet, trophic level and palaeoecology from bone stable isotope analysis: A comparative study of five red deer populations. Oecologia, 149: 12-21. [PDF]

T Coulson, JM Gaillard, M Festa-Bianchet (2005) Decomposing the variation in population growth into contributions from multiple demographic rates. Journal of Animal Ecology, 74: 789-801. [PDF]

DH Nussey, TH Clutton-Brock, SD Albon, J Pemberton, LEB Kruuk (2005) Constraints on plastic responses to climate variation in red deer. Biology Letters, 1: 457-460. [PDF]

DH Nussey, TH Clutton-Brock, DA Elston, SD Albon, LEB Kruuk (2005) Phenotypic plasticity in a maternal trait in red deer. Journal of Animal Ecology, 74: 387-396. [PDF]

DH Nussey, DW Coltman, T Coulson, LEB Kruuk, A Donald, SJ Morris, TH Clutton-Brock, J Pemberton (2005) Rapidly declining fine-scale spatial genetic structure in female red deer. Molecular Ecology, 14: 3395-3405. [PDF]

D Reby, K McComb, B Cargnelutti, C Darwin, WT Fitch, T Clutton-Brock (2005) Red deer stags use formants as assessment cues during intrasexual agonistic interactions. Proceedings of the Royal Society B-Biological Sciences, 272: 941-947. [PDF]

EA Catchpole, Y Fan, BJT Morgan, TH Clutton-Brock, T Coulson (2004) Sexual dimorphism, survival and dispersal in red deer. Journal of Agricultural Biological And Environmental Statistics, 9: 1-26. [PDF]

T Coulson, F Guinness, J Pemberton, T Clutton-Brock (2004) the demographic consequences of releasing a population of red deer from culling. Ecology, 85: 411-422. [PDF]

EJ Milner-Gulland, T Coulson, TH Clutton-Brock (2004) Sex differences and data quality as determinants of income from hunting red deer Cervus elaphus. Wildlife Biology, 10: 187-201. 

T Coulson, LEB Kruuk, G Tavecchia, JM Pemberton, TH Clutton-Brock (2003) Estimating selection on neonatal traits in red deer using elasticity path analysis. Evolution, 57: 2879-2892. [PDF]

LEB Kruuk, J Slate, JM Pemberton, TH Clutton-Brock (2003) Fluctuating asymmetry in a secondary sexual trait: No associations with individual fitness, environmental stress or inbreeding, and no heritability. Journal of Evolutionary Biology, 16: 101-113. [PDF]

D Reby, K McComb (2003) Anatomical constraints generate honesty: Acoustic cues to age and weight in the roars of red deer stags. Animal Behaviour, 65: 519-530. [PDF]

TH Clutton-Brock, TN Coulson, EJ Milner-Gulland, D Thomson, HM Armstrong (2002) Sex differences in emigration and mortality affect optimal management of deer populations. Nature, 415: 633-637. [PDF]

TH Clutton-Brock, T Coulson (2002) Comparative ungulate dynamics: the devil is in the detail. Philosophical Transactions of the Royal Society of London Series B-Biological Sciences, 357: 1285-1298. [PDF]

LEB Kruuk, J Slate, JM Pemberton, S Brotherstone, F Guinness, T Clutton-Brock (2002) Antler size in red deer: Heritability and selection but no evolution. Evolution, 56: 1683-1695. [PDF]

TC Marshall, DW Coltman, JM Pemberton, J Slate, JA Spalton, FE Guinness, JA Smith, JG Pilkington, TH Clutton-Brock (2002) Estimating the prevalence of inbreeding from incomplete pedigrees. Proceedings of the Royal Society of London Series B-Biological Sciences, 269: 1533-1539. [PDF]

J Slate, PM Visscher, S MacGregor, D Stevens, ML Tate, JM Pemberton (2002) A genome scan for quantitative trait loci in a wild population of red deer (Cervus elaphus). Genetics, 162: 1863-1873. [PDF]

J Slate, JM Pemberton (2002) Comparing molecular measures for detecting inbreeding depression. Journal of Evolutionary Biology, 15: 20-31. [PDF]

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Full Publication List

Books & Book Chapters

TH Clutton-Brock, FE Guinness, SD Albon (1982) Red deer: Behaviour and ecology of two sexes. University of Chicago Press, Chicago.

TH Clutton-Brock, SD Albon (1989) Red deer in the Highlands. BSP Professional Books, Oxford.

TH Clutton-Brock, SD Albon, FE Guinness (1988) Reproductive success in male and female red deer. In: Reproductive success (ed. Clutton-Brock TH), pp. 325-343. University of Chicago Press, Chicago.

SD Albon, TH Clutton-Brock (1988) Climate and the population dynamics of red deer in scotland. In: Ecological change in the uplands. (eds. Usher MB, Thompson DBA), pp. 93-106. Blackwell Scientific, Oxford.

SD Albon, TH Clutton-Brock, R Langvatn (1992) Cohort variation in reproduction and survival: Implications for population demography. In: The biology of deer (ed. Brown RD), pp. 15-21. Springer Verlag, Berlin.

JM Pemberton, JA Smith, TN Coulson, TC Marshall, J Slate, S Paterson, SD Albon, TH Clutton-Brock (1998) the maintenance of genetic polymorphism in small island populations: Large mammals in the hebrides. In: Evolution and islands (ed. Grant PR). Oxford University Press, Oxford.

JM Pemberton, DW Coltman, TN Coulson, J Slate (1999) Using microsatellites to measure the fitness consequences of inbreeding and outbreeding. In: Microsatellites: Evolution and applications (eds. Goldstein D, Schlotterer C). Oxford University Press, Oxford.


Papers

K Moyes, BJT Morgan, A Morris, SJ Morris, TH Clutton-Brock, T Coulson (2009) Exploring individual quality in a wild population of red deer. Journal of Animal Ecology 78: 406-413

DH Nussey, AJ Wilson, A Morris, J Pemberton, T Clutton-Brock, LEB Kruuk (2008) Testing for genetic trade-offs between early- and late-life reproduction in a wild red deer population. Proceedings of the Royal Society B-Biological Sciences 275: 745-750

KV Stopher, JM Pemberton,TH Clutton-Brock, T Coulson (2008) Individual differences, density dependence and offspring birth traits in a population of red deer. Proceedings of the Royal Society B-Biological Sciences 275: 2137-2145

T Coulson, S Tuljapurkar (2008) The dynamics of a quantitative trait in an age-structured population living in a variable environment. American Naturalist 172: 599-612.

PD McLoughlin, T Coulson, T Clutton-Brock (2008) Cross-generational effects of habitat and density on life history in red deer. Ecology 89: 3317-3326

DH Nussey, LEB Kruuk, A Morris, TH Clutton-Brock (2007) Environmental conditions in early life influence ageing rates in a wild population of red deer. Current Biology 17: R1000-R1001

AJ Wilson,  DH Nussey, JM Pemberton, JG Pilkington, A Morris, F Pelletier, TH Clutton-Brock, LEB Kruuk (2007) Evidence for a genetic basis of aging in two wild vertebrate populations. Current Biology 17: 2136-2142

K Foerster, T Coulson, BC Sheldon, JM Pemberton, TH Clutton-Brock, LEB Kruuk (2007) Sexually antagonistic genetic variation for fitness in red deer. Nature, 447: 1107-1110.

J Hone, TH Clutton-Brock (2007) Climate, food, density and wildlife population growth rate. Journal of Animal Ecology, 76: 361-367.

ST Kalinowski, ML Taper, TC Marshall (2007) Revising how the computer program CERVUS accommodates genotyping error increases success in paternity assignment. Molecular Ecology, 16: 1099-1106.

LEB Kruuk, JD Hadfield (2007) How to separate genetic and environmental causes of similarity between relatives. Journal of Evolutionary Biology, 20: 1890-1903.

DH Nussey, B Metherell, K Moyes, A Donald, FE Guinness, TH Clutton-Brock (2007) the relationship between tooth wear, habitat quality and late-life reproduction in a wild red deer population. Journal of Animal Ecology, 76: 402-412.

J Slate, JM Pemberton (2007) Admixture and patterns of linkage disequilibrium in a free-living vertebrate population. Journal of Evolutionary Biology, 20: 1415-1427.

K Csillery, T Johnson, D Beraldi, T Clutton-Brock, D Coltman, B Hansson, G Spong, JM Pemberton (2006) Performance of marker-based relatedness estimators in natural populations of outbred vertebrates. Genetics, 173: 2091-2101.

R Lande, S Engen, BE Saether, T Coulson (2006) Estimating density dependence from time series of population age structure. American Naturalist, 168: 76-87.

PD McLoughlin, MS Boyce, T Coulson, T Clutton-Brock (2006) Lifetime reproductive success and density-dependent, multi-variable resource selection. Proceedings of the Royal Society B-Biological Sciences, 273: 1449-1454.

K Moyes, T Coulson, BJT Morgan, A Donald, SJ Morris, TH Clutton-Brock (2006) Cumulative reproduction and survival costs in female red deer. Oikos, 115: 241-252.

DH Nussey, LEB Kruuk, A Donald, M Fowlie, TH Clutton-Brock (2006) the rate of senescence in maternal performance increases with early-life fecundity in red deer. Ecology Letters, 9: 1342-1350.

DH Nussey, J Pemberton, A Donald, LEB Kruuk (2006) Genetic consequences of human management in an introduced island population of red deer (Cervus elaphus). Heredity, 97: 56-65.

RE Stevens, AM Lister, REM Hedges (2006) Predicting diet, trophic level and palaeoecology from bone stable isotope analysis: A comparative study of five red deer populations. Oecologia, 149: 12-21.

T Coulson, JM Gaillard, M Festa-Bianchet (2005) Decomposing the variation in population growth into contributions from multiple demographic rates. Journal of Animal Ecology, 74: 789-801.

DH Nussey, TH Clutton-Brock, SD Albon, J Pemberton, LEB Kruuk (2005) Constraints on plastic responses to climate variation in red deer. Biology Letters, 1: 457-460.

DH Nussey, TH Clutton-Brock, DA Elston, SD Albon, LEB Kruuk (2005) Phenotypic plasticity in a maternal trait in red deer. Journal of Animal Ecology, 74: 387-396.

DH Nussey, DW Coltman, T Coulson, LEB Kruuk, A Donald, SJ Morris, TH Clutton-Brock, J Pemberton (2005) Rapidly declining fine-scale spatial genetic structure in female red deer. Molecular Ecology, 14: 3395-3405.

D Reby, K McComb, B Cargnelutti, C Darwin, WT Fitch, T Clutton-Brock (2005) Red deer stags use formants as assessment cues during intrasexual agonistic interactions. Proceedings of the Royal Society B-Biological Sciences, 272: 941-947.

EA Catchpole, Y Fan, BJT Morgan, TH Clutton-Brock, T Coulson (2004) Sexual dimorphism, survival and dispersal in red deer. Journal of Agricultural Biological And Environmental Statistics, 9: 1-26.

T Coulson, F Guinness, J Pemberton, T Clutton-Brock (2004) the demographic consequences of releasing a population of red deer from culling. Ecology, 85: 411-422.

EJ Milner-Gulland, T Coulson, TH Clutton-Brock (2004) Sex differences and data quality as determinants of income from hunting red deer Cervus elaphus. Wildlife Biology, 10: 187-201.

T Coulson, LEB Kruuk, G Tavecchia, JM Pemberton, TH Clutton-Brock (2003) Estimating selection on neonatal traits in red deer using elasticity path analysis. Evolution, 57: 2879-2892.

LEB Kruuk, J Slate, JM Pemberton, TH Clutton-Brock (2003) Fluctuating asymmetry in a secondary sexual trait: No associations with individual fitness, environmental stress or inbreeding, and no heritability. Journal of Evolutionary Biology, 16: 101-113.

D Reby, K McComb (2003) Anatomical constraints generate honesty: Acoustic cues to age and weight in the roars of red deer stags. Animal Behaviour, 65: 519-530.

TH Clutton-Brock, TN Coulson, EJ Milner-Gulland, D Thomson, HM Armstrong (2002) Sex differences in emigration and mortality affect optimal management of deer populations. Nature, 415: 633-637.

TH Clutton-Brock, T Coulson (2002) Comparative ungulate dynamics: the devil is in the detail. Philosophical Transactions of the Royal Society of London Series B-Biological Sciences, 357: 1285-1298.

LEB Kruuk, J Slate, JM Pemberton, S Brotherstone, F Guinness, T Clutton-Brock (2002) Antler size in red deer: Heritability and selection but no evolution. Evolution, 56: 1683-1695.

TC Marshall, DW Coltman, JM Pemberton, J Slate, JA Spalton, FE Guinness, JA Smith, JG Pilkington, TH Clutton-Brock (2002) Estimating the prevalence of inbreeding from incomplete pedigrees. Proceedings of the Royal Society of London Series B-Biological Sciences, 269: 1533-1539.

J Slate, PM Visscher, S MacGregor, D Stevens, ML Tate, JM Pemberton (2002) A genome scan for quantitative trait loci in a wild population of red deer (Cervus elaphus). Genetics, 162: 1863-1873.

J Slate, JM Pemberton (2002) Comparing molecular measures for detecting inbreeding depression. Journal of Evolutionary Biology, 15: 20-31.

L Conradt, IJ Gordon, TH Clutton-Brock, D Thomson, FE Guinness (2001) Could the indirect competition hypothesis explain inter-sexual site segregation in red deer (Cervus elaphus l.)? Journal of Zoology, 254: 185-193.

KT Schmidt, A Stien, SD Albon, FE Guinness (2001) Antler length of yearling red deer is determined by population density, weather and early life-history. Oecologia, 127: 191-197.

SD Albon, TN Coulson, D Brown, FE Guinness, JM Pemberton, TH Clutton-Brock (2000) Temporal changes in key factors and key age groups influencing the population dynamics of female red deer. Journal of Animal Ecology, 69: 1099-1110.

L Conradt, TH Clutton-Brock, FE Guinness (2000) Sex differences in weather sensitivity can cause habitat segregation: Red deer as an example. Animal Behaviour, 59: 1049-1060.

L Conradt (2000) Use of a seaweed habitat by red deer (Cervus elaphus l.). Journal of Zoology, 250: 541-549.

T Coulson, EJ Milner-Gulland, T Clutton-Brock (2000) the relative roles of density and climatic variation on population dynamics and fecundity rates in three contrasting ungulate species. Proceedings of the Royal Society of London Series B-Biological Sciences, 267: 1771-1779.

LEB Kruuk, TH Clutton-Brock, J Slate, JM Pemberton, S Brotherstone, FE Guinness (2000) Heritability of fitness in a wild mammal population. Proceedings of the National Academy of Sciences of the United States of America, 97: 698-703.

EJ Milner-Gulland, TN Coulson, TH Clutton-Brock (2000) On harvesting a structured ungulate population. Oikos, 88: 592-602.

J Slate, T Marshall, J Pemberton (2000) A retrospective assessment of the accuracy of the paternity inference program Cervus. Molecular Ecology, 9: 801-808.

J Slate, LEB Kruuk, TC Marshall, JM Pemberton, TH Clutton-Brock (2000) Inbreeding depression influences lifetime breeding success in a wild population of red deer (Cervus elaphus). Proceedings of the Royal Society of London Series B-Biological Sciences, 267: 1657-1662.

L Conradt (1999) Social segregation is not a consequence of habitat segregation in red deer and feral soay sheep. Animal Behaviour, 57: 1151-1157.

L Conradt, TH Clutton-Brock, D Thomson (1999) Habitat segregation in ungulates: Are males forced into suboptimal foraging habitats through indirect competition by females? Oecologia, 119: 367-377.

L Conradt, TH Clutton-Brock, FE Guinness (1999) the relationship between habitat choice and lifetime reproductive success in female red deer. Oecologia, 120: 218-224.

T Coulson, S Albon, J Slate, J Pemberton (1999) Microsatellite loci reveal sex-dependent responses to inbreeding and outbreeding in red deer calves. Evolution, 53: 1951-1960.

LEB Kruuk, TH Clutton-Brock, SD Albon, JM Pemberton, FE Guinness (1999) Population density affects sex ratio variation in red deer. Nature, 399: 459-461.

LEB Kruuk, TH Clutton-Brock, KE Rose, FE Guinness (1999) Early determinants of lifetime reproductive success differ between the sexes in red deer. Proceedings of the Royal Society of London Series B-Biological Sciences, 266: 1655-1661.

J Slate, JM Pemberton, PM Visscher (1999) Power to detect QTL in a free-living polygynous Ruminant population. Heredity, 83: 327-336.

L Conradt (1998) Measuring the degree of sexual segregation in group-living animals. Journal of Animal Ecology, 67: 217-226.

L Conradt (1998) Could asynchrony in activity between the sexes cause intersexual social segregation in ruminants? Proceedings of the Royal Society of London Series B-Biological Sciences, 265: 1359-1363.

TN Coulson, SD Albon, JM Pemberton, J Slate, FE Guinness, TH Clutton-Brock (1998) Genotype by environment interactions in winter survival in red deer. Journal of Animal Ecology, 67: 434-445.

TN Coulson, JM Pemberton, SD Albon, M Beaumont, TC Marshall, J Slate, FE Guinness, TH Clutton-Brock (1998) Microsatellites reveal heterosis in red deer. Proceedings of the Royal Society of London Series B-Biological Sciences, 265: 489-495.

TC Marshall, J Slate, LEB Kruuk, JM Pemberton (1998) Statistical confidence for likelihood-based paternity inference in natural populations. Molecular Ecology, 7: 639-655.

KE Rose, TH Clutton-Brock, FE Guinness (1998) Cohort variation in male survival and lifetime breeding success in red deer. Journal of Animal Ecology, 67: 979-986.

J Slate, DW Coltman, SJ Goodman, I MacLean, JM Pemberton, JL Williams (1998) Bovine microsatellite loci are highly conserved in red deer (Cervus elaphus), sika deer (Cervus nippon) and soay sheep (ovis aries). Animal Genetics, 29: 307-315.

TH Clutton-Brock, AW Illius, K Wilson, BT Grenfell, ADC MacColl, SD Albon (1997) Stability and instability in ungulate populations: An empirical analysis. American Naturalist, 149: 195-219.

TH Clutton-Brock, KE Rose, FE Guinness (1997) Density-related changes in sexual selection in red deer. Proceedings of the Royal Society of London Series B-Biological Sciences, 264: 1509-1516.

T Coulson, S Albon, F Guinness, J Pemberton, T CluttonBrock (1997) Population substructure, local density, and calf winter survival in red deer (Cervus elaphus). Ecology, 78: 852-863.

APF Flint, SD Albon, SI Jafar (1997) Blastocyst development and conceptus sex selection in red deer Cervus elaphus: Studies of a free-living population on the isle of Rum. General and Comparative Endocrinology, 106: 374-383.

APF Flint, SD Albon, ASI Loudon, J H.N. (1997) Behavioral dominance and corpus luteum function in red deer Cervus elaphus. Hormones and Behaviour, 31: 296-304.

R Langvatn, SD Albon, T Burkey, TH Clutton-Brock (1996) Climate, plant phenology and variation in age of first reproduction in a temperate herbivore. Journal of Animal Ecology, 65: 653-670.

JM Pemberton, JA Smith, TN Coulson, TC Marshall, J Slate, S Paterson, SD Albon, TH Clutton-Brock (1996) the maintenance of genetic polymorphism in small island populations: Large mammals in the hebrides. Philosophical Transactions of the Royal Society of London Series B-Biological Sciences, 351: 745-752.

TG Benton, A Grant, TH Clutton-Brock (1995) Does environmental stochasticity matter - analysis of red deer life-histories on Rum. Evolutionary Ecology, 9: 559-574.

JM Pemberton, J Slate, DR Bancroft, JA Barrett (1995) Nonamplifying alleles at microsatellite loci - a caution for parentage and population studies. Molecular Ecology, 4: 249-252.

TH Clutton-Brock, ME Lonergan (1994) Culling regimes and sex ratio biases in highland red deer. Journal of Applied Ecology, 31: 521-527.

SD Albon, HJ Staines, FE Guinness, TH Clutton-Brock (1992) Density-dependent changes in the spacing behavior of female kin in red deer. Journal of Animal Ecology, 61: 131-137.

JM Pemberton, SD Albon, FE Guinness, TH Clutton-Brock, GA Dover (1992) Behavioral estimates of male mating success tested by dna fingerprinting in a polygynous mammal. Behavioral Ecology, 3: 66-75.

JM Pemberton, SD Albon, FE Guinness, TH Clutton-Brock (1991) Countervailing selection in different fitness components in female red deer. Evolution, 45: 93-103.

M Gomendio, TH Clutton-Brock, SD Albon, FE Guinness, MJ Simpson (1990) Mammalian sex ratios and variation in costs of rearing sons and daughters. Nature, 343: 261-263.

TH Clutton-Brock, SD Albon, FE Guinness (1989) Fitness costs of gestation and lactation in wild mammals. Nature, 337: 260-262.

JM Pemberton, SD Albon, FE Guinness, TH Clutton-Brock, RJ Berry (1988) Genetic variation and juvenile survival in red deer. Evolution, 42: 921-934.

SD Albon, TH Clutton-Brock, FE Guinness (1987) Early development and population dynamics in red deer.2. Density-independent effects and cohort variation. Journal of Animal Ecology, 56: 69-81.

TH Clutton-Brock, M Major, SD Albon, FE Guinness (1987) Early development and population dynamics in red deer.1. Density-dependent effects on juvenile survival. Journal of Animal Ecology, 56: 53-67.

TH Clutton-Brock, SD Albon, FE Guinness (1987) Interactions between population-density and maternal characteristics affecting fecundity and juvenile survival in red deer. Journal of Animal Ecology, 56: 857-871.

TH Clutton-Brock, GR Iason, FE Guinness (1987) Sexual segregation and density-related changes in habitat use in male and female red deer (Cervus-elaphus). Journal of Zoology, 211: 275-289.

K McComb (1987) Roaring by red deer stags advances the date of estrus in hinds. Nature, 330: 648-649.

SD Albon, B Mitchell, BJ Huby, D Brown (1986) Fertility in female red deer (Cervus elaphus) - the effects of body-composition, age and reproductive status. Journal of Zoology, 209: 447-460.

TH Clutton-Brock, SD Albon, FE Guinness (1986) Great expectations - dominance, breeding success and offspring sex ratios in red deer. Animal Behaviour, 34: 460-471.

GR Iason, CD Duck, TH Clutton-Brock (1986) Grazing and reproductive success of red deer - the effect of local enrichment by gull colonies. Journal of Animal Ecology, 55: 507-515.

CR Thouless, FE Guinness (1986) Conflict between red deer hinds - the winner always wins. Animal Behaviour, 34: 1166-1171.

TH Clutton-Brock, M Major, FE Guinness (1985) Population regulation in male and female red deer. Journal of Animal Ecology, 54: 831-846.

TH Clutton-Brock, SD Albon, FE Guinness (1985) Parental investment and sex-differences in juvenile mortality in birds and mammals. Nature, 313: 131-133.

TH Clutton-Brock (1985) Reproductive success in red deer. Scientific American, 252: 86-.

GR Iason, FE Guinness (1985) Synchrony of estrus and conception in red deer (Cervus elaphus l). Animal Behaviour, 33: 1169-1174.

TH Clutton-Brock (1984) Reproductive effort and terminal investment in iteroparous animals. American Naturalist, 123: 212-229.

TH Clutton-Brock, SD Albon, FE Guinness (1984) Maternal dominance, breeding success and birth sex ratios in red deer. Nature, 308: 358-360.

TH Clutton-Brock, FE Guinness, SD Albon (1984) Individuals and populations - the effects of social-behavior on population-dynamics in deer. Proceedings of the Royal Society of Edinburgh Section B-Biological Sciences, 82: 275-290.

SD Albon, B Mitchell, BW Staines (1983) Fertility and body weight in female red deer - a density-dependent relationship. Journal of Animal Ecology, 52: 969-980.

SD Albon, FE Guinness, TH Clutton-Brock (1983) the influence of climatic variation on the birth weights of red deer (Cervus elaphus). Journal of Zoology, 200: 295-298.

TH Clutton-Brock, FE Guinness, SD Albon (1983) the costs of reproduction to red deer hinds. Journal of Animal Ecology, 52: 367-383.

TH Clutton-Brock, SD Albon (1983) Climatic variation and body weight of red deer. Journal of Wildlife Management, 47: 1197-1201.

T Clutton-Brock (1982) the red deer of Rhum. Natural History, 91: 42-.

TH Clutton-Brock (1982) the functions of antlers. Behaviour, 79: 108-.

TH Clutton-Brock, GR Iason, SD Albon, FE Guinness (1982) Effects of lactation on feeding behavior and habitat use in wild red deer hinds. Journal of Zoology, 198: 227-236.

TH Clutton-Brock, SD Albon (1982) Winter mortality in red deer (Cervus elaphus). Journal of Zoology, 198: 515-519.

TH Clutton-Brock, SD Albon, FE Guinness (1982) Competition between female relatives in a matrilocal mammal. Nature, 300: 178-180.

TH Clutton-Brock, SD Albon, FE Guinness (1981) Parental investment in male and female offspring in polygynous mammals. Nature, 289: 487-489.

RM Gibson, FE Guinness (1980) Differential reproduction among red deer (Cervus elaphus) stags on Rhum. Journal of Animal Ecology, 49: 199-208.

TH Clutton-Brock, SD Albon, RM Gibson, FE Guinness (1979) Logical stag - adaptive aspects of fighting in red deer (Cervus elaphus l). Animal Behaviour, 27: 211-225.

TH Clutton-Brock, SD Albon (1979) Roaring of red deer and the evolution of honest advertisement. Behaviour, 69: 145-.

FE Guinness, MJ Hall, RA Cockerill (1979) Mother-offspring association in red deer (Cervus elaphus l) on Rhum. Animal Behaviour, 27: 536-544.

FE Guinness, TH Clutton-Brock, SD Albon (1978) Factors affecting calf mortality in red deer (Cervus elaphus). Journal of Animal Ecology, 47: 817-.

FE Guinness, SD Albon, TH Clutton-Brock (1978) Factors affecting reproduction in red deer (Cervus-elaphus) hinds on Rhum. Journal of Reproduction And Fertility, 54: 325-334.

FE Guinness, RM Gibson, TH Clutton-Brock (1978) Calving times of red deer (Cervus-elaphus) on Rhum. Journal of Zoology, 185: 105-114.

TH Clutton-Brock, FE Guinness (1975) Behavior of red deer (Cervus elaphus l) at calving time. Behaviour, 55: 287-.

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