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Department of Zoology




  • 2006 – 2010: University of Southampton, Ph.D. Environmental Science. Thesis: The feasibility of reintroducing the burbot, Lota lota, to England. 
  • 2003 – 2004: University of East Anglia, MSc Applied Ecology and Conservation (Distinction)
  • 2000 – 2004: University of East Anglia, BSc Environmental Science


  • 2017 – to date: Research Associate, Conservation Science Group, Dept. Zoology, University of Cambridge, Cambridge (UK)
  • 2015 – 2016: Research Associate, Cardiff School of Biosciences, Cardiff University, Cardiff (UK)
  • 2011 – 2015: Postdoctoral Research Fellow, Oklahoma Cooperative Fish and Wildlife Research Unit, Stillwater, Oklahoma (USA)
  • 2010 – 2011: Postdoctoral Research Fellow, Faculty of Engineering and the Environment, University of Southampton, Southampton (UK)
  • 2004 – 2006: Ecological Appraisal Officer, Environment Agency, Litchfield & Nottingham (UK) 



Key publications: 

Worthington T., Echelle A.A., Perkin J.S., Mollenhauer R., Farless N., Dyer J.J., Longue, D. & Brewer S.K., (017). The emblematic minnows of the North American Great Plains: A synthesis of threats and conservation opportunities. Fish and Fisheries.

Brewer S.K., McManamay R.A., Miller A.D., Mollenhauer R., Worthington T. & Arsuffi, T. (2016) Advancing environmental flow science: developing frameworks for altered landscapes and integrating efforts across disciplines. Environmental Management, 58:175-192.  

Worthington T., Brewer S.K., Grabowski T.B. & Mueller J. (2014) Backcasting the decline of a vulnerable Great Plains reproductive ecotype: Identifying threats and conservation priorities. Global Change Biology, 20:89-102.

Kemp P.S., Worthington T. & Langford T.E.L. (2011) Quantifying the impact of reintroducing beaver on fish populations: a meta-analysis of the reviewed literature and survey of expert opinion. Fish and Fisheries, 13: 158-181.

Worthington T., Kemp P., Osborne P.E., Howes C. & Easton K. (2010) Former distribution and decline of the burbot (Lota lota) in the UK. Aquatic Conservation: Marine and Freshwater Ecosystems, 20: 371–377. 

Other publications: 

Brewer S.K., Worthington T., Mollenhauer R., Stewart D.R., McManamay R.A., Guertault L. & Moore D. (2018) Synthesizing models useful for ecohydrology and ecohydraulic approaches: An emphasis on integrating models to address complex research questions. Ecohydrology.

Musselman, W.C., Worthington T., Mouser, J., Williams, D. & Brewer S.K. (2017) Passive integrated transponder (PIT) tags: A review of current studies on warmwater fishes with notes on additional species. Journal of Fish and Wildlife Management, 8: 353-364.

Mueller J., Grabowski T.B., Brewer S.K., & Worthington T. (2017) Effects of temperature, dissolved solids, and suspended solids on the early life history stages of Arkansas River Shiner. Journal of Fish and Wildlife Management.

Worthington T., Zhang T., Logue D.R., Mittelstet A.R. & Brewer S.K. (2016) Landscape and flow metrics affecting the distribution of a federally-threatened fish: Improving management, model fit, and model transferability. Ecological Modelling, 342: 1-18. 

Bagwill A., Lovern M., Worthington T., Smith L. & McMurry S. (2016) Effects of water loss on New Mexico spadefoot toad (Spea multiplicata) development, spleen cellularity, and corticosterone levels. Journal of Experimental Zoology, 325: 548-561.

Worthington T., Van Houdt J.K.J., Hull J.M. & Kemp P. (2016) Thermal refugia and the survival of species in changing environments: new evidence from a nationally extinct freshwater fish. Ecology of Freshwater Fish.

Vowles A., Don A., Karageorgopoulos P., Worthington T. & Kemp P. (2015) Efficiency of a novel fish pass designed to mitigate for impeded upstream passage of juvenile European eels (Anguilla anguilla) at a model Crump weir. Fisheries Management and Ecology, 22: 307–316.)

Worthington T., Shaw P.J., Daffern J.R. & Langford T.E.L. (2015) The effects of a thermal discharge of the benthic macroinvertebrate community of the River Severn: implications for climate change. Hydrobiologia, 753: 81-95.

Dyer J.J., Worthington T. & Brewer S.K. (2015) Response of narrow-range crayfish to hyporheic water availability and excess sedimentation. Hydrobiologia, 747: 147-157.

Worthington T., Brewer S.K., Farless N., Grabowski T.B. & Gregory M.S. (2014) Interacting effects of discharge and channel morphology on transport of semibuoyant fish eggs in large, altered river systems. PLoS One, 9(5):e96599-e96608.

Worthington T., Brewer S.K., & Farless N. (2013) Spatial and temporal variation in efficiency of the Moore Egg Collector. North American Journal of Fisheries Management, 33: 1113-1118.)

Dyer J.J., Brewer S.K., Worthington T. & Bergey E.A. (2013) The influence of coarse-scale environmental features on current and future distributions of narrow-range endemic crayfish populations. Freshwater Biology, 58:1071-1088.

Worthington T., Brewer S.K., Grabowski T.B. & Mueller J. (2013) Sampling efficiency of the Moore egg collector. North American Journal of Fisheries Management, 33: 79-88.

Worthington T., Kemp P., Osborne P.E., Dillen A., Coeck J., Bunzel-Drüke M., Naura M., Gregory J. & Easton K. (2012) A spatial analytical GIS and RHS approach to selecting reintroduction sites for the reestablishment of the burbot in English rivers. Freshwater Biology, 57: 602-611.

Langford T.E.L., Worthington T., Shaw P.J., Kemp P.S., Ferguson A.J.D., Harding P. & Ottewell P. (2012) The Unnatural History of the River Trent: 50 Years of Ecological Recovery. In P.J Boon & P.J. Raven (Eds.) River Conservation and Management (pp. 261-272). John Wiley & Sons, Ltd.

Worthington T., Kemp P., Osborne P.E., Howes C. & Easton K. (2011) A review of the historical distribution and status of the burbot (Lota lota) in English rivers. Journal of Applied Ichthyology, 27 (Suppl. 1): 1–8. Worthington T., Kemp P., & Osborne P.E. (2011) Factors affecting the population viability of the burbot, Lota lota. Fisheries Management and Ecology, 18: 322–332.

Worthington T., Tisdale J., Kemp P., Williams I & Osborne P.E. (2010) Public and stakeholder attitudes to the reintroduction of the burbot, Lota lota. Fisheries Management and Ecology, 17: 465–472.

Worthington T., Vught I., De Charlroy D., Kemp P., Coeck J., Osborne P., Easton K. (2009) The re-introduction of the burbot to the United Kingdom and Flanders. In P.S. Soorae (Ed.), Global re-introduction perspectives: re-introduction case-studies from around the globe (pp. 26-29). Abu Dhabi: IUCN/SCC Re-introduction Specialist Group.

Research Associate

Contact Details

Room 3.01 David Attenborough Building